string: …lowfrequency loss of outer-hair cells and a small loss of apical and basal inner-hair cells (for review, see Willott, 1991; Saitoh et al., 1994; Gratton et al., 1996, 1997; Spongr et al., 1997; Ingham et al., 1999; Turner and Caspary, 2005) and auditory nerve fiber loss (Keithley et al., 1989,…