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Alkali metal | 666 | Production and isolation | Although sodium is less reactive than potassium, this process works because at such high temperatures potassium is more volatile than sodium and can easily be distilled off, so that the equilibrium shifts towards the right to produce more potassium gas and proceeds almost to completion. |
Alkali metal | 666 | Production and isolation | Metals like sodium are obtained by electrolysis of molten salts. Rb & Cs obtained mainly as by products of Li processing. To make pure caesium, ores of caesium and rubidium are crushed and heated to 650 °C with sodium metal, generating an alloy that can then be separated via a fractional distillation technique. Because metallic caesium is too reactive to handle, it is normally offered as caesium azide (CsN3). Caesium hydroxide is formed when caesium interacts aggressively with water and ice (CsOH). |
Alkali metal | 666 | Production and isolation | Rubidium is the 16th most prevalent element in the earth's crust; however, it is quite rare. Some minerals found in North America, South Africa, Russia, and Canada contain rubidium. Some potassium minerals (lepidolites, biotites, feldspar, carnallite) contain it, together with caesium. Pollucite, carnallite, leucite, and lepidolite are all minerals that contain rubidium. As a by-product of lithium extraction, it is commercially obtained from lepidolite. Rubidium is also found in potassium rocks and brines, which is a commercial supply. The majority of rubidium is now obtained as a byproduct of refining lithium. Rubidium is used in vacuum tubes as a getter, a material that combines with and removes trace gases from vacuum tubes. |
Alkali metal | 666 | Production and isolation | For several years in the 1950s and 1960s, a by-product of the potassium production called Alkarb was a main source for rubidium. Alkarb contained 21% rubidium while the rest was potassium and a small fraction of caesium. Today the largest producers of caesium, for example the Tanco Mine in Manitoba, Canada, produce rubidium as by-product from pollucite. Today, a common method for separating rubidium from potassium and caesium is the fractional crystallisation of a rubidium and caesium alum (Cs, Rb)Al(SO4)2·12H2O, which yields pure rubidium alum after approximately 30 recrystallisations. The limited applications and the lack of a mineral rich in rubidium limit the production of rubidium compounds to 2 to 4 tonnes per year. Caesium, however, is not produced from the above reaction. Instead, the mining of pollucite ore is the main method of obtaining pure caesium, extracted from the ore mainly by three methods: acid digestion, alkaline decomposition, and direct reduction. Both metals are produced as by-products of lithium production: after 1958, when interest in lithium's thermonuclear properties increased sharply, the production of rubidium and caesium also increased correspondingly. Pure rubidium and caesium metals are produced by reducing their chlorides with calcium metal at 750 °C and low pressure. |
Alkali metal | 666 | Production and isolation | As a result of its extreme rarity in nature, most francium is synthesised in the nuclear reaction Au + O → Fr + 5 n, yielding francium-209, francium-210, and francium-211. The greatest quantity of francium ever assembled to date is about 300,000 neutral atoms, which were synthesised using the nuclear reaction given above. When the only natural isotope francium-223 is specifically required, it is produced as the alpha daughter of actinium-227, itself produced synthetically from the neutron irradiation of natural radium-226, one of the daughters of natural uranium-238. |
Alkali metal | 666 | Applications | Lithium, sodium, and potassium have many applications, while rubidium and caesium are very useful in academic contexts but do not have many applications yet. Lithium is often used in lithium-ion batteries, and lithium oxide can help process silica. Lithium stearate is a thickener and can be used to make lubricating greases; it is produced from lithium hydroxide, which is also used to absorb carbon dioxide in space capsules and submarines. Lithium chloride is used as a brazing alloy for aluminium parts. Metallic lithium is used in alloys with magnesium and aluminium to give very tough and light alloys. |
Alkali metal | 666 | Applications | Sodium compounds have many applications, the most well-known being sodium chloride as table salt. Sodium salts of fatty acids are used as soap. Pure sodium metal also has many applications, including use in sodium-vapour lamps, which produce very efficient light compared to other types of lighting, and can help smooth the surface of other metals. Being a strong reducing agent, it is often used to reduce many other metals, such as titanium and zirconium, from their chlorides. Furthermore, it is very useful as a heat-exchange liquid in fast breeder nuclear reactors due to its low melting point, viscosity, and cross-section towards neutron absorption. |
Alkali metal | 666 | Applications | Potassium compounds are often used as fertilisers as potassium is an important element for plant nutrition. Potassium hydroxide is a very strong base, and is used to control the pH of various substances. Potassium nitrate and potassium permanganate are often used as powerful oxidising agents. Potassium superoxide is used in breathing masks, as it reacts with carbon dioxide to give potassium carbonate and oxygen gas. Pure potassium metal is not often used, but its alloys with sodium may substitute for pure sodium in fast breeder nuclear reactors. |
Alkali metal | 666 | Applications | Rubidium and caesium are often used in atomic clocks. Caesium atomic clocks are extraordinarily accurate; if a clock had been made at the time of the dinosaurs, it would be off by less than four seconds (after 80 million years). For that reason, caesium atoms are used as the definition of the second. Rubidium ions are often used in purple fireworks, and caesium is often used in drilling fluids in the petroleum industry. |
Alkali metal | 666 | Applications | Francium has no commercial applications, but because of francium's relatively simple atomic structure, among other things, it has been used in spectroscopy experiments, leading to more information regarding energy levels and the coupling constants between subatomic particles. Studies on the light emitted by laser-trapped francium-210 ions have provided accurate data on transitions between atomic energy levels, similar to those predicted by quantum theory. |
Alkali metal | 666 | Biological role and precautions | Pure alkali metals are dangerously reactive with air and water and must be kept away from heat, fire, oxidising agents, acids, most organic compounds, halocarbons, plastics, and moisture. They also react with carbon dioxide and carbon tetrachloride, so that normal fire extinguishers are counterproductive when used on alkali metal fires. Some Class D dry powder extinguishers designed for metal fires are effective, depriving the fire of oxygen and cooling the alkali metal. |
Alkali metal | 666 | Biological role and precautions | Experiments are usually conducted using only small quantities of a few grams in a fume hood. Small quantities of lithium may be disposed of by reaction with cool water, but the heavier alkali metals should be dissolved in the less reactive isopropanol. The alkali metals must be stored under mineral oil or an inert atmosphere. The inert atmosphere used may be argon or nitrogen gas, except for lithium, which reacts with nitrogen. Rubidium and caesium must be kept away from air, even under oil, because even a small amount of air diffused into the oil may trigger formation of the dangerously explosive peroxide; for the same reason, potassium should not be stored under oil in an oxygen-containing atmosphere for longer than 6 months. |
Alkali metal | 666 | Biological role and precautions | The bioinorganic chemistry of the alkali metal ions has been extensively reviewed. Solid state crystal structures have been determined for many complexes of alkali metal ions in small peptides, nucleic acid constituents, carbohydrates and ionophore complexes. |
Alkali metal | 666 | Biological role and precautions | Lithium naturally only occurs in traces in biological systems and has no known biological role, but does have effects on the body when ingested. Lithium carbonate is used as a mood stabiliser in psychiatry to treat bipolar disorder (manic-depression) in daily doses of about 0.5 to 2 grams, although there are side-effects. Excessive ingestion of lithium causes drowsiness, slurred speech and vomiting, among other symptoms, and poisons the central nervous system, which is dangerous as the required dosage of lithium to treat bipolar disorder is only slightly lower than the toxic dosage. Its biochemistry, the way it is handled by the human body and studies using rats and goats suggest that it is an essential trace element, although the natural biological function of lithium in humans has yet to be identified. |
Alkali metal | 666 | Biological role and precautions | Sodium and potassium occur in all known biological systems, generally functioning as electrolytes inside and outside cells. Sodium is an essential nutrient that regulates blood volume, blood pressure, osmotic equilibrium and pH; the minimum physiological requirement for sodium is 500 milligrams per day. Sodium chloride (also known as common salt) is the principal source of sodium in the diet, and is used as seasoning and preservative, such as for pickling and jerky; most of it comes from processed foods. The Dietary Reference Intake for sodium is 1.5 grams per day, but most people in the United States consume more than 2.3 grams per day, the minimum amount that promotes hypertension; this in turn causes 7.6 million premature deaths worldwide. |
Alkali metal | 666 | Biological role and precautions | Potassium is the major cation (positive ion) inside animal cells, while sodium is the major cation outside animal cells. The concentration differences of these charged particles causes a difference in electric potential between the inside and outside of cells, known as the membrane potential. The balance between potassium and sodium is maintained by ion transporter proteins in the cell membrane. The cell membrane potential created by potassium and sodium ions allows the cell to generate an action potential—a "spike" of electrical discharge. The ability of cells to produce electrical discharge is critical for body functions such as neurotransmission, muscle contraction, and heart function. Disruption of this balance may thus be fatal: for example, ingestion of large amounts of potassium compounds can lead to hyperkalemia strongly influencing the cardiovascular system. Potassium chloride is used in the United States for lethal injection executions. |
Alkali metal | 666 | Biological role and precautions | Due to their similar atomic radii, rubidium and caesium in the body mimic potassium and are taken up similarly. Rubidium has no known biological role, but may help stimulate metabolism, and, similarly to caesium, replace potassium in the body causing potassium deficiency. Partial substitution is quite possible and rather non-toxic: a 70 kg person contains on average 0.36 g of rubidium, and an increase in this value by 50 to 100 times did not show negative effects in test persons. Rats can survive up to 50% substitution of potassium by rubidium. Rubidium (and to a much lesser extent caesium) can function as temporary cures for hypokalemia; while rubidium can adequately physiologically substitute potassium in some systems, caesium is never able to do so. There is only very limited evidence in the form of deficiency symptoms for rubidium being possibly essential in goats; even if this is true, the trace amounts usually present in food are more than enough. |
Alkali metal | 666 | Biological role and precautions | Caesium compounds are rarely encountered by most people, but most caesium compounds are mildly toxic. Like rubidium, caesium tends to substitute potassium in the body, but is significantly larger and is therefore a poorer substitute. Excess caesium can lead to hypokalemia, arrythmia, and acute cardiac arrest, but such amounts would not ordinarily be encountered in natural sources. As such, caesium is not a major chemical environmental pollutant. The median lethal dose (LD50) value for caesium chloride in mice is 2.3 g per kilogram, which is comparable to the LD50 values of potassium chloride and sodium chloride. Caesium chloride has been promoted as an alternative cancer therapy, but has been linked to the deaths of over 50 patients, on whom it was used as part of a scientifically unvalidated cancer treatment. |
Alkali metal | 666 | Biological role and precautions | Radioisotopes of caesium require special precautions: the improper handling of caesium-137 gamma ray sources can lead to release of this radioisotope and radiation injuries. Perhaps the best-known case is the Goiânia accident of 1987, in which an improperly-disposed-of radiation therapy system from an abandoned clinic in the city of Goiânia, Brazil, was scavenged from a junkyard, and the glowing caesium salt sold to curious, uneducated buyers. This led to four deaths and serious injuries from radiation exposure. Together with caesium-134, iodine-131, and strontium-90, caesium-137 was among the isotopes distributed by the Chernobyl disaster which constitute the greatest risk to health. Radioisotopes of francium would presumably be dangerous as well due to their high decay energy and short half-life, but none have been produced in large enough amounts to pose any serious risk. |
Alphabet | 670 | An alphabet is a standardized set of written letters that represent particular spoken sounds in a language. Specifically, letters correspond to phonemes, the categories of sounds that can distinguish one word from another in a given language. Not all writing systems represent language in this way: a syllabary assigns symbols to spoken syllables, while logographic systems assign symbols to spoken words, morphemes, or other semantic units. |
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Alphabet | 670 | The first letters were invented in Ancient Egypt to aid writers already using Egyptian hieroglyphs, now referred to by lexicographers as the Egyptian uniliteral signs. This system was used until the 5th century AD, and fundamentally differed by adding pronunciation hints to existing hieroglyphs that had previously carried no pronunciation information. Later on, these phonemic symbols also became used to transcribe foreign words. The first fully phonemic script was the Proto-Sinaitic script, also descending from Egyptian hieroglyphics, which was later modified to create the Phoenician alphabet. The Phoenician system is considered the first true alphabet and is the ultimate ancestor of many modern scripts, including Arabic, Cyrillic, Greek, Hebrew, Latin, and possibly Brahmic. |
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Alphabet | 670 | Peter T. Daniels distinguishes true alphabets, which have letters representing both consonants and vowels, from both abugidas and abjads, which only have letters for consonants. Broadly, abjads lack vowel indicators altogether, while abugidas represent them with diacritics added to letters. In this narrower sense, the Greek alphabet was the first true alphabet, while the Phoenician alphabet it derived from was an abjad. |
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Alphabet | 670 | Alphabets are usually associated with a standard ordering of letters. This makes them useful for purposes of collation, which allows words to be sorted in a specific order, commonly known as the alphabetical order. It also means that their letters can be used as an alternative method of "numbering" ordered items, in such contexts as numbered lists and number placements. There are also names for letters in some languages. This is known as acrophony; It is present in some modern scripts, such as Greek, and many Semitic scripts, such as Arabic, Hebrew, and Syriac. It was used in some ancient alphabets, such as in Phoenician. However, this system is not present in all languages, such as the Latin alphabet, which adds a vowel after a character for each letter. Some systems also used to have this system but later on abandoned it for a system similar to Latin, such as Cyrillic. |
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Alphabet | 670 | Etymology | The English word alphabet came into Middle English from the Late Latin word alphabetum, which in turn originated in the Greek, ἀλφάβητος (alphábētos); it was made from the first two letters of the Greek alphabet, alpha (α) and beta (β). The names for the Greek letters, in turn, came from the first two letters of the Phoenician alphabet: aleph, the word for ox, and bet, the word for house. |
Alphabet | 670 | History | The Ancient Egyptian writing system had a set of some 24 hieroglyphs that are called uniliterals, which are glyphs that provide one sound. These glyphs were used as pronunciation guides for logograms, to write grammatical inflections, and, later, to transcribe loan words and foreign names. The script was used a fair amount in the 4th century CE. However, after pagan temples were closed down, it was forgotten in the 5th century until the discovery of the Rosetta Stone. There was also the Cuneiform script. The script was used to write several ancient languages. However, it was primarily used to write Sumerian. The last known use of the Cuneiform script was in 75 CE, after which the script fell out of use. |
Alphabet | 670 | History | In the Middle Bronze Age, an apparently "alphabetic" system known as the Proto-Sinaitic script appeared in Egyptian turquoise mines in the Sinai peninsula dated 1840 BCE, apparently left by Canaanite workers. Orly Goldwasser has connected the illiterate turquoise miner graffiti theory to the origin of the alphabet. In 1999, John and Deborah Darnell, American Egyptologists, discovered an earlier version of this first alphabet at the Wadi el-Hol valley in Egypt. The script dated to c. 1800 BCE and shows evidence of having been adapted from specific forms of Egyptian hieroglyphs that could be dated to c. 2000 BCE, strongly suggesting that the first alphabet had developed about that time. The script was based on letter appearances and names, believed to be based on Egyptian hieroglyphs. This script had no characters representing vowels. Originally, it probably was a syllabary—a script where syllables are represented with characters—with symbols that were not needed being removed. The best-attested Bronze Age alphabet is Ugaritic, invented in Ugarit (Syria) before the 15th century BCE. This was an alphabetic cuneiform script with 30 signs, including three that indicate the following vowel. This script was not used after the destruction of Ugarit in 1178 BCE. |
Alphabet | 670 | History | The Proto-Sinaitic script eventually developed into the Phoenician alphabet, conventionally called "Proto-Canaanite" before c. 1050 BCE. The oldest text in Phoenician script is an inscription on the sarcophagus of King Ahiram c. 1000 BCE. This script is the parent script of all western alphabets. By the tenth century BCE, two other forms distinguish themselves, Canaanite and Aramaic. The Aramaic gave rise to the Hebrew script. |
Alphabet | 670 | History | The South Arabian alphabet, a sister script to the Phoenician alphabet, is the script from which the Ge'ez alphabet, an abugida, a writing system where consonant-vowel sequences are written as units, which was used around the horn of Africa, descended. Vowel-less alphabets are called abjads, currently exemplified in others such as Arabic, Hebrew, and Syriac. The omission of vowels was not always a satisfactory solution due to the need of preserving sacred texts. "Weak" consonants are used to indicate vowels. These letters have a dual function since they can also be used as pure consonants. |
Alphabet | 670 | History | The Proto-Sinaitic script and the Ugaritic script were the first scripts with a limited number of signs instead of using many different signs for words, in contrast to the other widely used writing systems at the time, Cuneiform, Egyptian hieroglyphs, and Linear B. The Phoenician script was probably the first phonemic script, and it contained only about two dozen distinct letters, making it a script simple enough for traders to learn. Another advantage of the Phoenician alphabet was that it could write different languages since it recorded words phonemically. |
Alphabet | 670 | History | The Phoenician script was spread across the Mediterranean by the Phoenicians. The Greek Alphabet was the first alphabet in which vowels have independent letter forms separate from those of consonants. The Greeks chose letters representing sounds that did not exist in Phoenician to represent vowels. The syllabical Linear B, a script that was used by the Mycenaean Greeks from the 16th century BCE, had 87 symbols, including five vowels. In its early years, there were many variants of the Greek alphabet, causing many different alphabets to evolve from it. |
Alphabet | 670 | History | The Greek alphabet, in Euboean form, was carried over by Greek colonists to the Italian peninsula c. 800-600 BCE giving rise to many different alphabets used to write the Italic languages, like the Etruscan alphabet. One of these became the Latin alphabet, which spread across Europe as the Romans expanded their republic. After the fall of the Western Roman Empire, the alphabet survived in intellectual and religious works. It came to be used for the descendant languages of Latin (the Romance languages) and most of the other languages of western and central Europe. Today, it is the most widely used script in the world. |
Alphabet | 670 | History | The Etruscan alphabet remained nearly unchanged for several hundred years. Only evolving once the Etruscan language changed itself. The letters used for non-existent phonemes were dropped. Afterwards, however, the alphabet went through many different changes. The final classical form of Etruscan contained 20 letters. Four of them are vowels (a, e, i, and u). Six fewer letters than the earlier forms. The script in its classical form was used until the 1st century CE. The Etruscan language itself was not used in imperial Rome, but the script was used for religious texts. |
Alphabet | 670 | History | Some adaptations of the Latin alphabet have ligatures, a combination of two letters make one, such as æ in Danish and Icelandic and Ȣ in Algonquian; borrowings from other alphabets, such as the thorn þ in Old English and Icelandic, which came from the Futhark runes; and modified existing letters, such as the eth ð of Old English and Icelandic, which is a modified d. Other alphabets only use a subset of the Latin alphabet, such as Hawaiian and Italian, which uses the letters j, k, x, y, and w only in foreign words. |
Alphabet | 670 | History | Another notable script is Elder Futhark, believed to have evolved out of one of the Old Italic alphabets. Elder Futhark gave rise to other alphabets known collectively as the Runic alphabets. The Runic alphabets were used for Germanic languages from 100 CE to the late Middle Ages, being engraved on stone and jewelry, although inscriptions found on bone and wood occasionally appear. These alphabets have since been replaced with the Latin alphabet. The exception was for decorative use, where the runes remained in use until the 20th century. |
Alphabet | 670 | History | The Old Hungarian script was the writing system of the Hungarians. It was in use during the entire history of Hungary, albeit not as an official writing system. From the 19th century, it once again became more and more popular. |
Alphabet | 670 | History | The Glagolitic alphabet was the initial script of the liturgical language Old Church Slavonic and became, together with the Greek uncial script, the basis of the Cyrillic script. Cyrillic is one of the most widely used modern alphabetic scripts and is notable for its use in Slavic languages and also for other languages within the former Soviet Union. Cyrillic alphabets include Serbian, Macedonian, Bulgarian, Russian, Belarusian, and Ukrainian. The Glagolitic alphabet is believed to have been created by Saints Cyril and Methodius, while the Cyrillic alphabet was created by Clement of Ohrid, their disciple. They feature many letters that appear to have been borrowed from or influenced by Greek and Hebrew. |
Alphabet | 670 | History | Many phonetic scripts exist in Asia. The Arabic alphabet, Hebrew alphabet, Syriac alphabet, and other abjads of the Middle East are developments of the Aramaic alphabet. |
Alphabet | 670 | History | Most alphabetic scripts of India and Eastern Asia descend from the Brahmi script, believed to be a descendant of Aramaic. |
Alphabet | 670 | History | European alphabets, especially Latin and Cyrillic, have been adapted for many languages of Asia. Arabic is also widely used, sometimes as an abjad, as with Urdu and Persian, and sometimes as a complete alphabet, as with Kurdish and Uyghur. |
Alphabet | 670 | History | |
Alphabet | 670 | History | In Korea, Sejong the Great created the Hangul alphabet in 1443 CE. Hangul is a unique alphabet: it is a featural alphabet, where the design of many of the letters comes from a sound's place of articulation, like P looking like the widened mouth and L looking like the tongue pulled in. The creation of Hangul was planned by the government of the day, and it places individual letters in syllable clusters with equal dimensions, in the same way as Chinese characters. This change allows for mixed-script writing, where one syllable always takes up one type space no matter how many letters get stacked into building that one sound-block. |
Alphabet | 670 | History | Zhuyin, sometimes referred to as Bopomofo, is a semi-syllabary. It transcribes Mandarin phonetically in the Republic of China. After the later establishment of the People's Republic of China and its adoption of Hanyu Pinyin, the use of Zhuyin today is limited. However, it is still widely used in Taiwan. Zhuyin developed from a form of Chinese shorthand based on Chinese characters in the early 1900s and has elements of both an alphabet and a syllabary. Like an alphabet, the phonemes of syllable initials are represented by individual symbols, but like a syllabary, the phonemes of the syllable finals are not; each possible final (excluding the medial glide) has its own character, an example being luan written as ㄌㄨㄢ (l-u-an). The last symbol ㄢ takes place as the entire final -an. While Zhuyin is not a mainstream writing system, it is still often used in ways similar to a romanization system, for aiding pronunciation and as an input method for Chinese characters on computers and cellphones. |
Alphabet | 670 | Types | The term "alphabet" is used by linguists and paleographers in both a wide and a narrow sense. In a broader sense, an alphabet is a segmental script at the phoneme level—that is, it has separate glyphs for individual sounds and not for larger units such as syllables or words. In the narrower sense, some scholars distinguish "true" alphabets from two other types of segmental script, abjads, and abugidas. These three differ in how they treat vowels. Abjads have letters for consonants and leave most vowels unexpressed. Abugidas are also consonant-based but indicate vowels with diacritics, a systematic graphic modification of the consonants. The earliest known alphabet using this sense is the Wadi el-Hol script, believed to be an abjad. Its successor, Phoenician, is the ancestor of modern alphabets, including Arabic, Greek, Latin (via the Old Italic alphabet), Cyrillic (via the Greek alphabet), and Hebrew (via Aramaic). |
Alphabet | 670 | Types | Examples of present-day abjads are the Arabic and Hebrew scripts; true alphabets include Latin, Cyrillic, and Korean Hangul; and abugidas, used to write Tigrinya, Amharic, Hindi, and Thai. The Canadian Aboriginal syllabics are also an abugida, rather than a syllabary, as their name would imply, because each glyph stands for a consonant and is modified by rotation to represent the following vowel. In a true syllabary, each consonant-vowel combination gets represented by a separate glyph. |
Alphabet | 670 | Types | All three types may be augmented with syllabic glyphs. Ugaritic, for example, is essentially an abjad but has syllabic letters for /ʔa, ʔi, ʔu/ These are the only times that vowels are indicated. Coptic has a letter for /ti/. Devanagari is typically an abugida augmented with dedicated letters for initial vowels, though some traditions use अ as a zero consonant as the graphic base for such vowels. |
Alphabet | 670 | Types | The boundaries between the three types of segmental scripts are not always clear-cut. For example, Sorani Kurdish is written in the Arabic script, which, when used for other languages, is an abjad. In Kurdish, writing the vowels is mandatory, and whole letters are used, so the script is a true alphabet. Other languages may use a Semitic abjad with forced vowel diacritics, effectively making them abugidas. On the other hand, the Phagspa script of the Mongol Empire was based closely on the Tibetan abugida, but vowel marks are written after the preceding consonant rather than as diacritic marks. Although short a is not written, as in the Indic abugidas, The source of the term "abugida", namely the Ge'ez abugida now used for Amharic and Tigrinya, has assimilated into their consonant modifications. It is no longer systematic and must be learned as a syllabary rather than as a segmental script. Even more extreme, the Pahlavi abjad eventually became logographic. |
Alphabet | 670 | Types | Thus the primary categorisation of alphabets reflects how they treat vowels. For tonal languages, further classification can be based on their treatment of tone. Though names do not yet exist to distinguish the various types. Some alphabets disregard tone entirely, especially when it does not carry a heavy functional load, as in Somali and many other languages of Africa and the Americas. Most commonly, tones are indicated by diacritics, which is how vowels are treated in abugidas, which is the case for Vietnamese (a true alphabet) and Thai (an abugida). In Thai, the tone is determined primarily by a consonant, with diacritics for disambiguation. In the Pollard script, an abugida, vowels are indicated by diacritics. The placing of the diacritic relative to the consonant is modified to indicate the tone. More rarely, a script may have separate letters for tones, as is the case for Hmong and Zhuang. For many, regardless of whether letters or diacritics get used, the most common tone is not marked, just as the most common vowel is not marked in Indic abugidas. In Zhuyin, not only is one of the tones unmarked; but there is a diacritic to indicate a lack of tone, like the virama of Indic. |
Alphabet | 670 | Alphabetical order | Alphabets often come to be associated with a standard ordering of their letters; this is for collation—namely, for listing words and other items in alphabetical order. |
Alphabet | 670 | Alphabetical order | The basic ordering of the Latin alphabet (A B C D E F G H I J K L M N O P Q R S T U V W X Y Z), which derives from the Northwest Semitic "Abgad" order, is already well established. Although, languages using this alphabet have different conventions for their treatment of modified letters (such as the French é, à, and ô) and certain combinations of letters (multigraphs). In French, these are not considered to be additional letters for collation. However, in Icelandic, the accented letters such as á, í, and ö are considered distinct letters representing different vowel sounds from sounds represented by their unaccented counterparts. In Spanish, ñ is considered a separate letter, but accented vowels such as á and é are not. The ll and ch were also formerly considered single letters and sorted separately after l and c, but in 1994, the tenth congress of the Association of Spanish Language Academies changed the collating order so that ll came to be sorted between lk and lm in the dictionary and ch came to be sorted between cg and ci; those digraphs were still formally designated as letters, but in 2010 the Real Academia Española changed it, so they are no longer considered letters at all. |
Alphabet | 670 | Alphabetical order | In German, words starting with sch- (which spells the German phoneme /ʃ/) are inserted between words with initial sca- and sci- (all incidentally loanwords) instead of appearing after the initial sz, as though it were a single letter, which contrasts several languages such as Albanian, in which dh-, ë-, gj-, ll-, rr-, th-, xh-, and zh-, which all represent phonemes and considered separate single letters, would follow the letters d, e, g, l, n, r, t, x, and z, respectively, as well as Hungarian and Welsh. Further, German words with an umlaut get collated ignoring the umlaut as—contrary to Turkish, which adopted the graphemes ö and ü, and where a word like tüfek would come after tuz, in the dictionary. An exception is the German telephone directory, where umlauts are sorted like ä=ae since names such as Jäger also appear with the spelling Jaeger and are not distinguished in the spoken language. |
Alphabet | 670 | Alphabetical order | The Danish and Norwegian alphabets end with æ—ø—å, whereas the Swedish conventionally put å—ä—ö at the end. However, æ phonetically corresponds with ä, as does ø and ö. |
Alphabet | 670 | Alphabetical order | It is unknown whether the earliest alphabets had a defined sequence. Some alphabets today, such as the Hanuno'o script, are learned one letter at a time, in no particular order, and are not used for collation where a definite order is required. However, a dozen Ugaritic tablets from the fourteenth century BCE preserve the alphabet in two sequences. One, the ABCDE order later used in Phoenician, has continued with minor changes in Hebrew, Greek, Armenian, Gothic, Cyrillic, and Latin; the other, HMĦLQ, was used in southern Arabia and is preserved today in Ethiopic. Both orders have therefore been stable for at least 3000 years. |
Alphabet | 670 | Alphabetical order | Runic used an unrelated Futhark sequence, which got simplified later on. Arabic uses usually uses its sequence, although Arabic retains the traditional abjadi order, which is used for numbers. |
Alphabet | 670 | Alphabetical order | The Brahmic family of alphabets used in India uses a unique order based on phonology: The letters are arranged according to how and where the sounds get produced in the mouth. This organization is present in Southeast Asia, Tibet, Korean hangul, and even Japanese kana, which is not an alphabet. |
Alphabet | 670 | Acrophony | In Phoenician, each letter got associated with a word that begins with that sound. This is called acrophony and is continuously used to varying degrees in Samaritan, Aramaic, Syriac, Hebrew, Greek, and Arabic. |
Alphabet | 670 | Acrophony | Acrophony got abandoned in Latin. It referred to the letters by adding a vowel (usually "e", sometimes "a", or "u") before or after the consonant. Two exceptions were Y and Z, which were borrowed from the Greek alphabet rather than Etruscan. They were known as Y Graeca "Greek Y" and zeta (from Greek)—this discrepancy was inherited by many European languages, as in the term zed for Z in all forms of English, other than American English. Over time names sometimes shifted or were added, as in double U for W, or "double V" in French, the English name for Y, and the American zee for Z. Comparing them in English and French gives a clear reflection of the Great Vowel Shift: A, B, C, and D are pronounced /eɪ, biː, siː, diː/ in today's English, but in contemporary French they are /a, be, se, de/. The French names (from which the English names got derived) preserve the qualities of the English vowels before the Great Vowel Shift. By contrast, the names of F, L, M, N, and S (/ɛf, ɛl, ɛm, ɛn, ɛs/) remain the same in both languages because "short" vowels were largely unaffected by the Shift. |
Alphabet | 670 | Acrophony | In Cyrillic, originally, acrophony was present using Slavic words. The first three words going, azŭ, buky, vědě, with the Cyrillic collation order being, А, Б, В. However, this was later abandoned in favor of a system similar to Latin. |
Alphabet | 670 | Orthography and pronunciation | When an alphabet is adopted or developed to represent a given language, an orthography generally comes into being, providing rules for spelling words, following the principle on which alphabets get based. These rules will map letters of the alphabet to the phonemes of the spoken language. In a perfectly phonemic orthography, there would be a consistent one-to-one correspondence between the letters and the phonemes so that a writer could predict the spelling of a word given its pronunciation, and a speaker would always know the pronunciation of a word given its spelling, and vice versa. However, this ideal is usually never achieved in practice. Languages can come close to it, such as Spanish and Finnish. others, such as English, deviate from it to a much larger degree. |
Alphabet | 670 | Orthography and pronunciation | The pronunciation of a language often evolves independently of its writing system. Writing systems have been borrowed for languages the orthography was not initially made to use. The degree to which letters of an alphabet correspond to phonemes of a language varies. |
Alphabet | 670 | Orthography and pronunciation | Languages may fail to achieve a one-to-one correspondence between letters and sounds in any of several ways: |
Alphabet | 670 | Orthography and pronunciation | National languages sometimes elect to address the problem of dialects by associating the alphabet with the national standard. Some national languages like Finnish, Armenian, Turkish, Russian, Serbo-Croatian (Serbian, Croatian, and Bosnian), and Bulgarian have a very regular spelling system with nearly one-to-one correspondence between letters and phonemes. Similarly, the Italian verb corresponding to 'spell (out),' compitare, is unknown to many Italians because spelling is usually trivial, as Italian spelling is highly phonemic. In standard Spanish, one can tell the pronunciation of a word from its spelling, but not vice versa, as phonemes sometimes can be represented in more than one way, but a given letter is consistently pronounced. French using silent letters, nasal vowels, and elision, may seem to lack much correspondence between the spelling and pronunciation. However, its rules on pronunciation, though complex, are consistent and predictable with a fair degree of accuracy. |
Alphabet | 670 | Orthography and pronunciation | At the other extreme are languages such as English, where pronunciations mostly have to be memorized as they do not correspond to the spelling consistently. For English, this is because the Great Vowel Shift occurred after the orthography got established and because English has acquired a large number of loanwords at different times, retaining their original spelling at varying levels. However, even English has general, albeit complex, rules that predict pronunciation from spelling. Rules like this are usually successful. However, rules to predict spelling from pronunciation have a higher failure rate. |
Alphabet | 670 | Orthography and pronunciation | Sometimes, countries have the written language undergo a spelling reform to realign the writing with the contemporary spoken language. These can range from simple spelling changes and word forms to switching the entire writing system. For example, Turkey switched from the Arabic alphabet to a Latin-based Turkish alphabet, and when Kazakh changed from an Arabic script to a Cyrillic script due to the Soviet Union's influence, and in 2021, it made a transition to the Latin alphabet, similar to Turkish. The Cyrillic script used to be official in Uzbekistan and Turkmenistan before they all switched to the Latin alphabet, including Uzbekistan that is having a reform of the alphabet to use diacritics on the letters that are marked by apostrophes and the letters that are digraphs. |
Alphabet | 670 | Orthography and pronunciation | The standard system of symbols used by linguists to represent sounds in any language, independently of orthography, is called the International Phonetic Alphabet. |
Atomic number | 673 | The atomic number or nuclear charge number (symbol Z) of a chemical element is the charge number of an atomic nucleus. For ordinary nuclei composed of protons and neutrons, this is equal to the proton number (np) or the number of protons found in the nucleus of every atom of that element. The atomic number can be used to uniquely identify ordinary chemical elements. In an ordinary uncharged atom, the atomic number is also equal to the number of electrons. |
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Atomic number | 673 | For an ordinary atom which contains protons, neutrons and electrons, the sum of the atomic number Z and the neutron number N gives the atom's atomic mass number A. Since protons and neutrons have approximately the same mass (and the mass of the electrons is negligible for many purposes) and the mass defect of the nucleon binding is always small compared to the nucleon mass, the atomic mass of any atom, when expressed in daltons (making a quantity called the "relative isotopic mass"), is within 1% of the whole number A. |
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Atomic number | 673 | Atoms with the same atomic number but different neutron numbers, and hence different mass numbers, are known as isotopes. A little more than three-quarters of naturally occurring elements exist as a mixture of isotopes (see monoisotopic elements), and the average isotopic mass of an isotopic mixture for an element (called the relative atomic mass) in a defined environment on Earth determines the element's standard atomic weight. Historically, it was these atomic weights of elements (in comparison to hydrogen) that were the quantities measurable by chemists in the 19th century. |
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Atomic number | 673 | The conventional symbol Z comes from the German word Zahl 'number', which, before the modern synthesis of ideas from chemistry and physics, merely denoted an element's numerical place in the periodic table, whose order was then approximately, but not completely, consistent with the order of the elements by atomic weights. Only after 1915, with the suggestion and evidence that this Z number was also the nuclear charge and a physical characteristic of atoms, did the word Atomzahl (and its English equivalent atomic number) come into common use in this context. |
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Atomic number | 673 | The rules above do not always apply to exotic atoms which contain short-lived elementary particles other than protons, neutrons and electrons. |
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Atomic number | 673 | History | Loosely speaking, the existence or construction of a periodic table of elements creates an ordering of the elements, and so they can be numbered in order. |
Atomic number | 673 | History | Dmitri Mendeleev claimed that he arranged his first periodic tables (first published on March 6, 1869) in order of atomic weight ("Atomgewicht"). However, in consideration of the elements' observed chemical properties, he changed the order slightly and placed tellurium (atomic weight 127.6) ahead of iodine (atomic weight 126.9). This placement is consistent with the modern practice of ordering the elements by proton number, Z, but that number was not known or suspected at the time. |
Atomic number | 673 | History | A simple numbering based on periodic table position was never entirely satisfactory. In addition to the case of iodine and tellurium, several other pairs of elements (such as argon and potassium, cobalt and nickel) were later shown to have nearly identical or reversed atomic weights, thus requiring their placement in the periodic table to be determined by their chemical properties. However the gradual identification of more and more chemically similar lanthanide elements, whose atomic number was not obvious, led to inconsistency and uncertainty in the periodic numbering of elements at least from lutetium (element 71) onward (hafnium was not known at this time). |
Atomic number | 673 | History | In 1911, Ernest Rutherford gave a model of the atom in which a central nucleus held most of the atom's mass and a positive charge which, in units of the electron's charge, was to be approximately equal to half of the atom's atomic weight, expressed in numbers of hydrogen atoms. This central charge would thus be approximately half the atomic weight (though it was almost 25% different from the atomic number of gold (Z = 79, A = 197), the single element from which Rutherford made his guess). Nevertheless, in spite of Rutherford's estimation that gold had a central charge of about 100 (but was element Z = 79 on the periodic table), a month after Rutherford's paper appeared, Antonius van den Broek first formally suggested that the central charge and number of electrons in an atom were exactly equal to its place in the periodic table (also known as element number, atomic number, and symbolized Z). This eventually proved to be the case. |
Atomic number | 673 | History | The experimental position improved dramatically after research by Henry Moseley in 1913. Moseley, after discussions with Bohr who was at the same lab (and who had used Van den Broek's hypothesis in his Bohr model of the atom), decided to test Van den Broek's and Bohr's hypothesis directly, by seeing if spectral lines emitted from excited atoms fitted the Bohr theory's postulation that the frequency of the spectral lines be proportional to the square of Z. |
Atomic number | 673 | History | To do this, Moseley measured the wavelengths of the innermost photon transitions (K and L lines) produced by the elements from aluminium (Z = 13) to gold (Z = 79) used as a series of movable anodic targets inside an x-ray tube. The square root of the frequency of these photons (x-rays) increased from one target to the next in an arithmetic progression. This led to the conclusion (Moseley's law) that the atomic number does closely correspond (with an offset of one unit for K-lines, in Moseley's work) to the calculated electric charge of the nucleus, i.e. the element number Z. Among other things, Moseley demonstrated that the lanthanide series (from lanthanum to lutetium inclusive) must have 15 members—no fewer and no more—which was far from obvious from known chemistry at that time. |
Atomic number | 673 | History | After Moseley's death in 1915, the atomic numbers of all known elements from hydrogen to uranium (Z = 92) were examined by his method. There were seven elements (with Z < 92) which were not found and therefore identified as still undiscovered, corresponding to atomic numbers 43, 61, 72, 75, 85, 87 and 91. From 1918 to 1947, all seven of these missing elements were discovered. By this time, the first four transuranium elements had also been discovered, so that the periodic table was complete with no gaps as far as curium (Z = 96). |
Atomic number | 673 | History | In 1915, the reason for nuclear charge being quantized in units of Z, which were now recognized to be the same as the element number, was not understood. An old idea called Prout's hypothesis had postulated that the elements were all made of residues (or "protyles") of the lightest element hydrogen, which in the Bohr-Rutherford model had a single electron and a nuclear charge of one. However, as early as 1907, Rutherford and Thomas Royds had shown that alpha particles, which had a charge of +2, were the nuclei of helium atoms, which had a mass four times that of hydrogen, not two times. If Prout's hypothesis were true, something had to be neutralizing some of the charge of the hydrogen nuclei present in the nuclei of heavier atoms. |
Atomic number | 673 | History | In 1917, Rutherford succeeded in generating hydrogen nuclei from a nuclear reaction between alpha particles and nitrogen gas, and believed he had proven Prout's law. He called the new heavy nuclear particles protons in 1920 (alternate names being proutons and protyles). It had been immediately apparent from the work of Moseley that the nuclei of heavy atoms have more than twice as much mass as would be expected from their being made of hydrogen nuclei, and thus there was required a hypothesis for the neutralization of the extra protons presumed present in all heavy nuclei. A helium nucleus was presumed to be composed of four protons plus two "nuclear electrons" (electrons bound inside the nucleus) to cancel two of the charges. At the other end of the periodic table, a nucleus of gold with a mass 197 times that of hydrogen was thought to contain 118 nuclear electrons in the nucleus to give it a residual charge of +79, consistent with its atomic number. |
Atomic number | 673 | History | All consideration of nuclear electrons ended with James Chadwick's discovery of the neutron in 1932. An atom of gold now was seen as containing 118 neutrons rather than 118 nuclear electrons, and its positive nuclear charge now was realized to come entirely from a content of 79 protons. Since Moseley had previously shown that the atomic number Z of an element equals this positive charge, it was now clear that Z is identical to the number of protons of its nuclei. |
Atomic number | 673 | Chemical properties | Each element has a specific set of chemical properties as a consequence of the number of electrons present in the neutral atom, which is Z (the atomic number). The configuration of these electrons follows from the principles of quantum mechanics. The number of electrons in each element's electron shells, particularly the outermost valence shell, is the primary factor in determining its chemical bonding behavior. Hence, it is the atomic number alone that determines the chemical properties of an element; and it is for this reason that an element can be defined as consisting of any mixture of atoms with a given atomic number. |
Atomic number | 673 | New elements | The quest for new elements is usually described using atomic numbers. As of 2023, all elements with atomic numbers 1 to 118 have been observed. Synthesis of new elements is accomplished by bombarding target atoms of heavy elements with ions, such that the sum of the atomic numbers of the target and ion elements equals the atomic number of the element being created. In general, the half-life of a nuclide becomes shorter as atomic number increases, though undiscovered nuclides with certain "magic" numbers of protons and neutrons may have relatively longer half-lives and comprise an island of stability. |
Atomic number | 673 | New elements | A hypothetical element composed only of neutrons has also been proposed and would have atomic number 0. |
Anatomy | 674 | Anatomy (from Ancient Greek ἀνατομή (anatomḗ) 'dissection') is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having its beginnings in prehistoric times. Anatomy is inherently tied to developmental biology, embryology, comparative anatomy, evolutionary biology, and phylogeny, as these are the processes by which anatomy is generated, both over immediate and long-term timescales. Anatomy and physiology, which study the structure and function of organisms and their parts respectively, make a natural pair of related disciplines, and are often studied together. Human anatomy is one of the essential basic sciences that are applied in medicine. |
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Anatomy | 674 | Anatomy is a complex and dynamic field that is constantly evolving as new discoveries are made. In recent years, there has been a significant increase in the use of advanced imaging techniques, such as MRI and CT scans, which allow for more detailed and accurate visualizations of the body's structures. |
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Anatomy | 674 | The discipline of anatomy is divided into macroscopic and microscopic parts. Macroscopic anatomy, or gross anatomy, is the examination of an animal's body parts using unaided eyesight. Gross anatomy also includes the branch of superficial anatomy. Microscopic anatomy involves the use of optical instruments in the study of the tissues of various structures, known as histology, and also in the study of cells. |
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Anatomy | 674 | The history of anatomy is characterized by a progressive understanding of the functions of the organs and structures of the human body. Methods have also improved dramatically, advancing from the examination of animals by dissection of carcasses and cadavers (corpses) to 20th-century medical imaging techniques, including X-ray, ultrasound, and magnetic resonance imaging. |
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Anatomy | 674 | Etymology and definition | Derived from the Greek ἀνατομή anatomē "dissection" (from ἀνατέμνω anatémnō "I cut up, cut open" from ἀνά aná "up", and τέμνω témnō "I cut"), anatomy is the scientific study of the structure of organisms including their systems, organs and tissues. It includes the appearance and position of the various parts, the materials from which they are composed, and their relationships with other parts. Anatomy is quite distinct from physiology and biochemistry, which deal respectively with the functions of those parts and the chemical processes involved. For example, an anatomist is concerned with the shape, size, position, structure, blood supply and innervation of an organ such as the liver; while a physiologist is interested in the production of bile, the role of the liver in nutrition and the regulation of bodily functions. |
Anatomy | 674 | Etymology and definition | The discipline of anatomy can be subdivided into a number of branches, including gross or macroscopic anatomy and microscopic anatomy. Gross anatomy is the study of structures large enough to be seen with the naked eye, and also includes superficial anatomy or surface anatomy, the study by sight of the external body features. Microscopic anatomy is the study of structures on a microscopic scale, along with histology (the study of tissues), and embryology (the study of an organism in its immature condition). Regional anatomy is the study of the interrelationships of all of the structures in a specific body region, such as the abdomen. In contrast, systemic anatomy is the study of the structures that make up a discrete body system—that is, a group of structures that work together to perform a unique body function, such as the digestive system. |
Anatomy | 674 | Etymology and definition | Anatomy can be studied using both invasive and non-invasive methods with the goal of obtaining information about the structure and organization of organs and systems. Methods used include dissection, in which a body is opened and its organs studied, and endoscopy, in which a video camera-equipped instrument is inserted through a small incision in the body wall and used to explore the internal organs and other structures. Angiography using X-rays or magnetic resonance angiography are methods to visualize blood vessels. |
Anatomy | 674 | Etymology and definition | The term "anatomy" is commonly taken to refer to human anatomy. However, substantially similar structures and tissues are found throughout the rest of the animal kingdom, and the term also includes the anatomy of other animals. The term zootomy is also sometimes used to specifically refer to non-human animals. The structure and tissues of plants are of a dissimilar nature and they are studied in plant anatomy. |
Anatomy | 674 | Animal tissues | The kingdom Animalia contains multicellular organisms that are heterotrophic and motile (although some have secondarily adopted a sessile lifestyle). Most animals have bodies differentiated into separate tissues and these animals are also known as eumetazoans. They have an internal digestive chamber, with one or two openings; the gametes are produced in multicellular sex organs, and the zygotes include a blastula stage in their embryonic development. Metazoans do not include the sponges, which have undifferentiated cells. |
Anatomy | 674 | Animal tissues | Unlike plant cells, animal cells have neither a cell wall nor chloroplasts. Vacuoles, when present, are more in number and much smaller than those in the plant cell. The body tissues are composed of numerous types of cells, including those found in muscles, nerves and skin. Each typically has a cell membrane formed of phospholipids, cytoplasm and a nucleus. All of the different cells of an animal are derived from the embryonic germ layers. Those simpler invertebrates which are formed from two germ layers of ectoderm and endoderm are called diploblastic and the more developed animals whose structures and organs are formed from three germ layers are called triploblastic. All of a triploblastic animal's tissues and organs are derived from the three germ layers of the embryo, the ectoderm, mesoderm and endoderm. |
Anatomy | 674 | Animal tissues | Animal tissues can be grouped into four basic types: connective, epithelial, muscle and nervous tissue. |
Anatomy | 674 | Animal tissues | Connective tissues are fibrous and made up of cells scattered among inorganic material called the extracellular matrix. Connective tissue gives shape to organs and holds them in place. The main types are loose connective tissue, adipose tissue, fibrous connective tissue, cartilage and bone. The extracellular matrix contains proteins, the chief and most abundant of which is collagen. Collagen plays a major part in organizing and maintaining tissues. The matrix can be modified to form a skeleton to support or protect the body. An exoskeleton is a thickened, rigid cuticle which is stiffened by mineralization, as in crustaceans or by the cross-linking of its proteins as in insects. An endoskeleton is internal and present in all developed animals, as well as in many of those less developed. |
Anatomy | 674 | Animal tissues | Epithelial tissue is composed of closely packed cells, bound to each other by cell adhesion molecules, with little intercellular space. Epithelial cells can be squamous (flat), cuboidal or columnar and rest on a basal lamina, the upper layer of the basement membrane, the lower layer is the reticular lamina lying next to the connective tissue in the extracellular matrix secreted by the epithelial cells. There are many different types of epithelium, modified to suit a particular function. In the respiratory tract there is a type of ciliated epithelial lining; in the small intestine there are microvilli on the epithelial lining and in the large intestine there are intestinal villi. Skin consists of an outer layer of keratinized stratified squamous epithelium that covers the exterior of the vertebrate body. Keratinocytes make up to 95% of the cells in the skin. The epithelial cells on the external surface of the body typically secrete an extracellular matrix in the form of a cuticle. In simple animals this may just be a coat of glycoproteins. In more advanced animals, many glands are formed of epithelial cells. |
Anatomy | 674 | Animal tissues | Muscle cells (myocytes) form the active contractile tissue of the body. Muscle tissue functions to produce force and cause motion, either locomotion or movement within internal organs. Muscle is formed of contractile filaments and is separated into three main types; smooth muscle, skeletal muscle and cardiac muscle. Smooth muscle has no striations when examined microscopically. It contracts slowly but maintains contractibility over a wide range of stretch lengths. It is found in such organs as sea anemone tentacles and the body wall of sea cucumbers. Skeletal muscle contracts rapidly but has a limited range of extension. It is found in the movement of appendages and jaws. Obliquely striated muscle is intermediate between the other two. The filaments are staggered and this is the type of muscle found in earthworms that can extend slowly or make rapid contractions. In higher animals striated muscles occur in bundles attached to bone to provide movement and are often arranged in antagonistic sets. Smooth muscle is found in the walls of the uterus, bladder, intestines, stomach, oesophagus, respiratory airways, and blood vessels. Cardiac muscle is found only in the heart, allowing it to contract and pump blood round the body. |
Anatomy | 674 | Animal tissues | Nervous tissue is composed of many nerve cells known as neurons which transmit information. In some slow-moving radially symmetrical marine animals such as ctenophores and cnidarians (including sea anemones and jellyfish), the nerves form a nerve net, but in most animals they are organized longitudinally into bundles. In simple animals, receptor neurons in the body wall cause a local reaction to a stimulus. In more complex animals, specialized receptor cells such as chemoreceptors and photoreceptors are found in groups and send messages along neural networks to other parts of the organism. Neurons can be connected together in ganglia. In higher animals, specialized receptors are the basis of sense organs and there is a central nervous system (brain and spinal cord) and a peripheral nervous system. The latter consists of sensory nerves that transmit information from sense organs and motor nerves that influence target organs. The peripheral nervous system is divided into the somatic nervous system which conveys sensation and controls voluntary muscle, and the autonomic nervous system which involuntarily controls smooth muscle, certain glands and internal organs, including the stomach. |
Anatomy | 674 | Vertebrate anatomy | All vertebrates have a similar basic body plan and at some point in their lives, mostly in the embryonic stage, share the major chordate characteristics: a stiffening rod, the notochord; a dorsal hollow tube of nervous material, the neural tube; pharyngeal arches; and a tail posterior to the anus. The spinal cord is protected by the vertebral column and is above the notochord, and the gastrointestinal tract is below it. Nervous tissue is derived from the ectoderm, connective tissues are derived from mesoderm, and gut is derived from the endoderm. At the posterior end is a tail which continues the spinal cord and vertebrae but not the gut. The mouth is found at the anterior end of the animal, and the anus at the base of the tail. The defining characteristic of a vertebrate is the vertebral column, formed in the development of the segmented series of vertebrae. In most vertebrates the notochord becomes the nucleus pulposus of the intervertebral discs. However, a few vertebrates, such as the sturgeon and the coelacanth, retain the notochord into adulthood. Jawed vertebrates are typified by paired appendages, fins or legs, which may be secondarily lost. The limbs of vertebrates are considered to be homologous because the same underlying skeletal structure was inherited from their last common ancestor. This is one of the arguments put forward by Charles Darwin to support his theory of evolution. |
Anatomy | 674 | Vertebrate anatomy | The body of a fish is divided into a head, trunk and tail, although the divisions between the three are not always externally visible. The skeleton, which forms the support structure inside the fish, is either made of cartilage, in cartilaginous fish, or bone in bony fish. The main skeletal element is the vertebral column, composed of articulating vertebrae which are lightweight yet strong. The ribs attach to the spine and there are no limbs or limb girdles. The main external features of the fish, the fins, are composed of either bony or soft spines called rays, which with the exception of the caudal fins, have no direct connection with the spine. They are supported by the muscles which compose the main part of the trunk. The heart has two chambers and pumps the blood through the respiratory surfaces of the gills and on round the body in a single circulatory loop. The eyes are adapted for seeing underwater and have only local vision. There is an inner ear but no external or middle ear. Low frequency vibrations are detected by the lateral line system of sense organs that run along the length of the sides of fish, and these respond to nearby movements and to changes in water pressure. |
Anatomy | 674 | Vertebrate anatomy | Sharks and rays are basal fish with numerous primitive anatomical features similar to those of ancient fish, including skeletons composed of cartilage. Their bodies tend to be dorso-ventrally flattened, they usually have five pairs of gill slits and a large mouth set on the underside of the head. The dermis is covered with separate dermal placoid scales. They have a cloaca into which the urinary and genital passages open, but not a swim bladder. Cartilaginous fish produce a small number of large, yolky eggs. Some species are ovoviviparous and the young develop internally but others are oviparous and the larvae develop externally in egg cases. |
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